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Telos (/ˈtɛ.lɒs/; Greek: τέλος, translit. télos, lit. "end, 'purpose', or 'goal'") is a term used by philosopher Aristotle to refer to the final cause of a natural organ or entity, or of human art. Telos is the root of the modern term teleology, the study of purposiveness or of objects with a view to their aims, purposes, or intentions. Teleology is central in Aristotle's work on plant and animal biology, and human ethics, through his theory of the four causes. Aristotle's notion that everything has a telos also gave rise to epistemology.